Theoretical studies predict that the stability of an ecosystem is negatively correlated with its complexity, measured by the number of interacting species. On the other hand, empirical evidence indicates that food webs are highly interconnected.
Food webs fall into two classes with clearly distinct stability properties. In one of them stability is negatively correlated with complexity, and in the other group stability is positively correlated. The positive or negative relationship is related only to the topological structure of the food web. It is independent of the number of connections, strengths of predator-prey interactions or number of species.